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MotacillaFlavaKeulemans. upper left Motacilla borealis Sund Billberg Sund Billberg Sund Billberg Sund SaviSavi Sund SaviSavi Sund SaviSavi Greyheaded Wagtail upper row and left center Ashyheaded Wagtail right center and lower row Catalogue of the birds in the British Museum archive org Vol Plate VII creator John Gerrard Keulemans